Sivapardus Temporal range: | |
---|---|
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Carnivora |
Suborder: | Feliformia |
Family: | Felidae |
Subfamily: | Felinae |
Genus: | † Sivapardus Bakr, 1969 |
Type species | |
Sivapardus punjabiensis Bakr, 1969 | |
Sar-Dhok locality (Punjab, Pakistan) |
Sivapardus is an extinct, little-known genus of feline with only one species assigned to it, Sivapardus punjabiensis. It was described in 1969 by the paleontologist Abu Bakr based on a partial mandible from the Upper Siwaliks in Pakistan; the locality it was found at is estimated to be from the Late Pliocene to Early Pleistocene. S. punjabiensis was a large cat with a short and broad snout that may have lived on open grasslands.
The type and only specimen, U.Z. No. 67/22 (part of the Punjab University Zoology Department Fossil Collection), was collected in 1967 from the Sar Dhok locality in the Pabbi Hills west of Kharain, Gujrat, in western Pakistan. It was described and assigned by Abu Bakr to the new genus and species Sivapardus punjabiensis in 1969. No etymology was given for either name. [1]
The type specimen is a right mandibular ramus with several teeth: the fourth premolar and first molar are damaged but present, but only the roots of the third premolar and the canine tooth remain, and the third incisor is represented only by its tooth socket. The third incisor was situated a little in front of the canine tooth, which itself seems to have been relatively very large. The diastema behind the canine tooth was short. The shape of the third premolar, based on the roots, is typical of cats. [1]
The fourth premolar, though damaged, resembles most felines in having three cusps in a row; the main cusp was likely slightly higher than the paraconid of the molar (similar to Panthera, not preserved in Sivapanthera), and a prominent posterior cingulum (a shelf at the base of the tooth) is located behind the posterior (rear) cusp (also similar to both Panthera and Sivapanthera); there is indication of a cingulum-like structure on the left and right of the anterior (front) cusp, but Bakr dismissed this as having any taxonomic value due to its small size. The first molar also retained enough of its structure for description: relative to the premolars, the anterior end is inclined towards the inside (like Panthera and unlike Acinonyx); the protoconid was longer and a cusp taller than the paraconid, with a deep valley between the two cusps; there is only faint indication of a talonid and no metaconid at all. The molar overall is longer than the fourth premolar, akin to Sivapanthera and unlike Panthera. [1]
Bakr described the species as similar to Sivapanthera in general proportions, with the major difference being the masseteric fossa (the depression where the masseter muscle attaches to the jaw), which in Sivapanthera specimens extended to at least the hind end of the first molar, but in Sivapardus punjabiensis ended abruptly well before the first molar, and the end of the fossa was well-defined and deep; this feature also set it apart from Panthera and Acinonyx . The distinct shape of the masseteric fossa was given as the diagnostic feature of the genus and species. [1]
Based on U.Z. No. 67/22, Bakr described Sivapardus punjabiensis as a cat with a short and broad snout similar to that of the cheetah-like Sivapanthera, larger in size than a leopard but smaller than a lion. [1]
Bakr classified Sivapardus as a feline, a member of the subfamily Felinae. [1]
The Sar-Dhok locality is estimated to be of Late Pliocene to late Early Pleistocene age, and may have been a savannah-like habitat, an open grasslands in a semi-arid climate, with denser growth along riverbanks. Other fauna known from the three Pabbi Hills localities (Sar-Dhok, Panjan Sher Shahana, and Kurla Sharif) are primarily herbivores and include proboscideans such as Elephas hysudricus , Elephas planifrons , and several species of Stegodon ; the bovids Boselaphus namadicus , Proamphibos kashmiricus , Hemibos triquetricornis , Bos acutifrons , Bubalus palaeindicus , Bubalus platyceros , Kobus porrecticornis , Sivacobus patulicornis , Antilope sp., Sivatragus bohlini , Damalops and an indeterminate caprine; the cervids Metacervocerus punjabiensis and Rucervus ; the giraffid Sivatherium giganteum ; the hippopotamid Hexaprotodon sivalensis ; rhinocerotids including Rhinoceros sivalensis , Rhinoceros sondaicus ; and the equid Equus sivalensis . [2]
In mammalian oral anatomy, the canine teeth, also called cuspids, dog teeth, eye teeth, vampire teeth, or vampire fangs, are the relatively long, pointed teeth. In the context of the upper jaw, they are also known as fangs. They can appear more flattened however, causing them to resemble incisors and leading them to be called incisiform. They developed and are used primarily for firmly holding food in order to tear it apart, and occasionally as weapons. They are often the largest teeth in a mammal's mouth. Individuals of most species that develop them normally have four, two in the upper jaw and two in the lower, separated within each jaw by incisors; humans and dogs are examples. In most species, canines are the anterior-most teeth in the maxillary bone. The four canines in humans are the two upper maxillary canines and the two lower mandibular canines. They are specially prominent in dogs (Canidae), hence the name.
Gigantopithecus is an extinct genus of ape from roughly 2 million to 350,000 years ago during the Early to Middle Pleistocene of southern China, represented by one species, Gigantopithecus blacki. Potential identifications have also been made in Thailand, Vietnam, and Indonesia. The first remains of Gigantopithecus, two third molar teeth, were identified in a drugstore by anthropologist Ralph von Koenigswald in 1935, who subsequently described the ape. In 1956, the first mandible and more than 1,000 teeth were found in Liucheng, and numerous more remains have since been found in at least 16 sites. Only teeth and four mandibles are known currently, and other skeletal elements were likely consumed by porcupines before they could fossilise. Gigantopithecus was once argued to be a hominin, a member of the human line, but it is now thought to be closely allied with orangutans, classified in the subfamily Ponginae.
Diatomys is an extinct rodent genus known from Miocene deposits in China, Japan, Pakistan, and Thailand. The fossil range is from the late Early Miocene to the Middle Miocene.
Dental anatomy is a field of anatomy dedicated to the study of human tooth structures. The development, appearance, and classification of teeth fall within its purview. Tooth formation begins before birth, and the teeth's eventual morphology is dictated during this time. Dental anatomy is also a taxonomical science: it is concerned with the naming of teeth and the structures of which they are made, this information serving a practical purpose in dental treatment.
Gobiconodon is an extinct genus of carnivorous mammals belonging to the family Gobiconodontidae. Undisputed records of Gobiconodon are restricted to the Early Cretaceous of Asia and North America, but isolated teeth attributed to the genus have also been described from formations in England and Morocco dating as far back as the Middle Jurassic. Species of Gobiconodon varied considerably in size, with G. ostromi, one of the larger species, being around the size of a modern Virginia opossum. Like other gobiconodontids, it possessed several speciations towards carnivory, such as shearing molariform teeth, large canine-like incisors and powerful jaw and forelimb musculature, indicating that it probably fed on vertebrate prey. Unusually among predatory mammals and other eutriconodonts, the lower canines were vestigial, with the first lower incisor pair having become massive and canine-like. Like the larger Repenomamus there might be some evidence of scavenging.
Asiavorator is an extinct genus of civet-like carnivoran belonging in the family Stenoplesictidae. It was endemic to Asia and lived during the Eocene and Oligocene epochs.
Hipposideros besaoka is an extinct bat from Madagascar in the genus Hipposideros. It is known from numerous jaws and teeth, which were collected in a cave at Anjohibe in 1996 and described as a new species in 2007. The site where H. besaoka was found is at most 10,000 years old; other parts of the cave have yielded H. commersoni, a living species of Hipposideros from Madagascar, and some material that is distinct from both species. H. besaoka was larger than H. commersoni, making it the largest insectivorous bat of Madagascar, and had broader molars and a more robust lower jaw. As usual in Hipposideros, the second upper premolar is small and displaced from the toothrow, and the second lower premolar is large.
Triaenops goodmani is an extinct bat from Madagascar in the genus Triaenops. It is known from three lower jaws collected in a cave at Anjohibe in 1996, and described as a new species in 2007. The material is at most 10,000 years old. A bat humerus from the same site could not be identified as either T. goodmani or the living T. menamena. T. goodmani is identifiable as a member of Triaenops or the related genus Paratriaenops by a number of features of the teeth, such as the single-cusped, canine-like fourth premolar and the presence of a gap between the entoconid and hypoconulid cusps on the first two molars. T. goodmani is larger than the living species of Triaenops and Paratriaenops on Madagascar, and on the first molar the protoconid cusp is only slightly higher than the hypoconid, not much higher as in the other species.
Miniopterus tao is a fossil bat in the genus Miniopterus from the Pleistocene of Zhoukoudian in China. It is known from a number of mandibles, which were initially identified as the living species Miniopterus schreibersii in 1963 before being recognized as a separate species, M. tao, in 1986. Miniopterus tao is larger than living M. schreibersii and has more closely spaced lower premolars and more robust talonids on the lower molars. The back part of the mandible is relatively low and on it, the coronoid and condyloid processes are about equally high. The average length of the mandible is 12.0 mm.
Lagrivea is a fossil genus of squirrel from the Middle Miocene of France. The single species, L. vireti, is known from three mandibles and two isolated teeth. All come from the fissure filling of La Grive L5, part of the La Grive-Saint-Alban complex in Saint-Alban-de-Roche, southeastern France. Lagrivea was a large tree squirrel with flat lower incisors and a large, triangular fourth lower premolar (p4). Each of the four cheekteeth bears a deep basin in the middle of the crown. The m3 is about rectangular in shape, but rounded at the back. Although m1 and m2 have two roots, m3 has three.
Dermotherium is a genus of fossil mammals closely related to the living colugos, a small group of gliding mammals from Southeast Asia. Two species are recognized: D. major from the Late Eocene of Thailand, based on a single fragment of the lower jaw, and D. chimaera from the Late Oligocene of Thailand, known from three fragments of the lower jaw and two isolated upper molars. In addition, a single isolated upper molar from the Early Oligocene of Pakistan has been tentatively assigned to D. chimaera. All sites where fossils of Dermotherium have been found were probably forested environments and the fossil species were probably forest dwellers like living colugos, but whether they had the gliding adaptations of the living species is unknown.
Indraloris is a fossil primate from the Miocene of India and Pakistan in the family Sivaladapidae. Two species are now recognized: I. himalayensis from Haritalyangar, India and I. kamlialensis from the Pothohar Plateau, Pakistan. Other material from the Potwar Plateau may represent an additional, unnamed species. Body mass estimates range from about 2 kg (4.4 lb) for the smaller I. kamlialensis to over 4 kg (8.8 lb) for the larger I. himalayensis.
Sivaladapis is a genus of adapiform primate that lived in Asia during the middle Miocene.
Hemimachairodus is an extinct genus of machairodontine (sabre-toothed) cat with only one species, Hemimachairodus zwierzyckii, known from fossils from the Pleistocene of Java.
Enhydriodon is an extinct genus of mustelids known from Africa, Pakistan, and India that lived from the late Miocene to the early Pleistocene. It contains 9 confirmed species, 2 debated species, and at least a few other undescribed species from Africa. The genus belongs to the tribe Enhydriodontini in the otter subfamily Lutrinae. Enhydriodon means “otter tooth” in Ancient Greek and is a reference to its dentition rather than to the Enhydra genus, which includes the modern sea otter and its two prehistoric relatives.
Taowu is an extinct genus of machairodonts, a type of saber-toothed cat. It lived during the Early Pleistocene about 2.5 million years ago in East Asia. So far, only one skull is known, found in northern China. Based on this, a relatively small representative of the saber-toothed cats can be reconstructed, which only reached the size of a present-day leopard. In its dentition characteristics, it mediates between phylogenetic older forms such as Amphimachairodus and younger members such as Homotherium. The genus was scientifically described in 2022, but the find material was recovered as early as the 1930s.
Cokotherium is an extinct genus of eutherian mammal from the Early Cretaceous of China. It includes a single species, Cokotherium jiufotangensis, known from a single partial skeleton, missing a portion of the hindlimbs and tail. It was recovered from the Jiufotang Formation, the upper part of the fossiliferous Jehol biota. The generic name of Cokotherium honors the nickname of the late paleontologist Chuan-Kui Li, a specialist on the Jiufotang Formation. The specific name refers to the formation in question. Cokotherium is one of the youngest and most well-preserved Early Cretaceous eutherians, illustrating an array of transitional conditions between Early Cretaceous and Late Cretaceous members of Eutheria.
Sivasmilus is a fossil genus of barbourofelid containing only a single species, Sivasmilus copei. It is known from only a single specimen, a partial mandible collected from the Chinji Formation in the Lower Siwaliks in Pakistan, estimated to be from the Miocene. The fossil was originally described in 1915 when it was assigned to the fossil feline Sivaelurus chinjiensis, but was used as the basis of a new genus and species in 1929 by Hungarian paleontologist Miklós Kretzoi. Sivasmilus copei was a relatively small, cat-like animal.
Lonchocyon is an extinct genus of arctoid carnivorans, with possible affinities to amphicyonids or hemicyonine bears. It contains a single species, Lonchocyon qiui, known from a single left mandible discovered at the fossil-bearing locality Haerhada at the base of the Baron Sog Formation, which is located in Inner Mongolia, China, and dates to the late Eocene. This taxon is notable for its large size in comparison to other arctoid carnivorans of the Eocene epoch, and for its hypercarnivorous adaptions, most notably its large canine and strongly reduced premolars. The genus name is a combination of Greek lonch, meaning spear and referencing the spear-like paraconid on its lower carnassial, and cyon, meaning dog. The specific name honours Professor Zhan-Xiang Qiu.
Pachypanthera is an extinct genus of pantherine felid that was recovered from the Late Miocene-aged Khorat sand pits in northeastern Thailand. It contains a single species, Pachypanthera piriyai, named and described in 2023.