Cheilanthes | |
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Cheilanthes parryi | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Division: | Polypodiophyta |
Class: | Polypodiopsida |
Order: | Polypodiales |
Family: | Pteridaceae |
Subfamily: | Cheilanthoideae |
Genus: | Cheilanthes Sw. [1] |
Type species | |
Cheilanthes micropteris Sw. | |
Species | |
See text | |
Synonyms | |
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Cheilanthes, commonly known as lip ferns, [2] is a genus of about 180 species of rock-dwelling ferns with a cosmopolitan distribution in warm, dry, rocky regions, often growing in small crevices high up on cliffs. Most are small, sturdy and evergreen. The leaves, often densely covered in trichomes, spring directly from the rootstocks. Many of them are desert ferns, curling up during dry times and reviving with the coming of moisture. At the ends of veins sporangia, or spore-bearing structures, are protected by leaf margins, which curl over them.
The genus name is derived from the Greek words χεῖλος (cheilos), meaning "lip," and ἄνθος (anthos), meaning "flower." [3]
Cheilanthes as traditionally circumscribed is now known to be highly paraphyletic, comprising at least four generically separate groups. The type species, C. micropteris , is most closely allied to the genera Aleuritopteris and Sinopteris (Schuettpelz et al.). In the early 21st century, many species, principally from the New World, were moved into the new genus Gaga and the revived genus Myriopteris . Further work remains to be done to render Cheilanthes monophyletic. Members of many other cheilanthoid genera have at times been given names in the genus.
The circumscription of the genus was uncertain as of January 2020 [update] . The Checklist of Ferns and Lycophytes of the World lists species in several distinct groups. [4]
Cheilanthes sensu stricto contains about 20 species from North and tropical Central and South America: [5]
Cheilanthes sensu lato contains about 50 species that molecular phylogenetic studies suggest do not form a clade with the core Cheilanthes species, falling into separate clades, but for which no placement outside Cheilanthes was available as of November 2019 [update] : [6]
Pteris (brake) is a genus of about 300 species of ferns in the subfamily Pteridoideae of the family Pteridaceae. They are native to tropical and subtropical regions, southward to New Zealand, Australia, and South Africa, north to Japan and North America. 78 species are found in China. Some species of Pteris have considerable economic and ecological value, such as Pteris multifida, Pteris ensiformis, Pteris vittata can be used for ornamental purposes; as a hyperaccumulator, Pteris multifida and Pteris vittata can be used to control soil pollution.
Cyathea is a genus of tree ferns, the type genus of the fern order Cyatheales.
Pellaea is a genus of ferns in the Cheilanthoideae subfamily of the Pteridaceae. The genus name is derived from the Greek word πελλος (pellos), meaning "dark," and refers to the brown stems. Many members of the genus are commonly known as cliffbrakes. They primarily grow in rocky habitats, including moist rocky canyons, slopes, and bluffs.
Notholaena, cloak fern, is a genus of ferns in the Cheilanthoideae subfamily of the Pteridaceae. Ferns of this genus are mostly epipetric or occurring in coarse, gravelly soils, and are most abundant and diverse in the mountain ranges of warm arid or semiarid regions. They typically have a creeping or erect rhizome and leaves that are pinnatifid to pinnate-pinnatifid with marginal sori protected by a false indusium formed from the reflexed margin of the leaf. Members of Notholaena also have a coating of whitish or yellowish farina on the surfaces of the leaves. The farina is often limited to the abaxial (lower) leaf surface, but may occur on the adaxial (upper) leaf surface as well. Members of the related Pentagramma genus have a similar lower leaf-surface farina.
Alsophila is a genus of tree ferns in the family Cyatheaceae. It has also been considered to be a section in the subgenus Cyathea of the genus Cyathea.
Dennstaedtia is a mostly tropical and subtropical group of ferns described as a genus in 1801. Hayscented fern, or cup ferns, are common names for some species in this genus. It includes 58 species native to the tropical Americas, Madagascar, southern and eastern Asia, Australia, New Guinea, and the Pacific Islands.
Pleopeltis is a genus of ferns in the family Polypodiaceae, subfamily Polypodioideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). The genus widely distributed in tropical regions of the world, and also north into temperate regions in eastern North America and eastern Asia. Several species are known by the common name scaly polypody and resurrection fern.
Campyloneurum is a genus of ferns in the family Polypodiaceae, subfamily Polypodioideae, according to the Pteridophyte Phylogeny Group classification of 2016 (PPG I). They are commonly known as strap ferns.
Tectaria is a genus of fern in the family Tectariaceae, according to the Pteridophyte Phylogeny Group classification of 2016. Halberd fern is a common name for species in this genus.
Argyrochosma is a genus of ferns known commonly as false cloak ferns. The genus is included in the Cheilanthoideae subfamily of the Pteridaceae. Species now in this genus were previously treated as members of related genera Notholaena or Pellaea but were segregated into their own genus in 1987. These ferns, of which there are about 20 species, are mostly native to the Americas, from North to South and including the Caribbean, while one species, A. connectens, is known from Sichuan, China. They are commonly found growing in cracks between rocks. Their leaves are generally shorter than 40 cm (16 in) and have rounded bluish or grayish green segments. Often the lower surface of the segments is coated in a white dust, and the sporangia contain brown spores.
Christella is a genus of around 70-80 species of ferns in the subfamily Thelypteridoideae of the family Thelypteridaceae in the Pteridophyte Phylogeny Group classification of 2016. Other sources sink Christella into a very broadly defined genus Thelypteris. The genus was named after Konrad H. Christ, a Swiss botanist. The distribution of these plants is mostly in the tropics and sub tropical areas. An Australian example is C. dentata.
Doryopteris is a genus of ferns in the subfamily Cheilanthoideae of the family Pteridaceae.
Adiantopsis is a genus of ferns in the subfamily Cheilanthoideae of the family Pteridaceae.
Pityrogramma, the silverback ferns, or goldback ferns, is a fern genus in the subfamily Pteridoideae of the family Pteridaceae.
Amauropelta is a genus of ferns in the family Thelypteridaceae, subfamily Thelypteridoideae, in the Pteridophyte Phylogeny Group classification of 2016. Other sources sink the genus into a very broadly defined genus Thelypteris.
Goniopteris is a genus of ferns in the family Thelypteridaceae, subfamily Thelypteridoideae, in the Pteridophyte Phylogeny Group classification of 2016. Other sources sink Goniopteris into a very broadly defined genus Thelypteris.
Stigmatopteris is a genus of ferns in the family Dryopteridaceae, subfamily Polybotryoideae, in the Pteridophyte Phylogeny Group classification of 2016.