Nyctaginaceae | |
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Garden Four O'Clock ( Mirabilis jalapa ) flowers | |
Scientific classification | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Clade: | Angiosperms |
Clade: | Eudicots |
Order: | Caryophyllales |
Family: | Nyctaginaceae Juss. [1] |
Tribes | |
Synonyms | |
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Nyctaginaceae, the four o'clock family, is a family of around 33 genera and 290 species of flowering plants, widely distributed in tropical and subtropical regions, with a few representatives in temperate regions. The family has a distinctive fruit type called an accessory fruit or anthocarp, and many genera have extremely large (>100 μm) pollen grains.
The family has been almost universally recognized by plant taxonomists. The APG II system (2003; unchanged from the APG system of 1998), assigns it to the order Caryophyllales in the clade core eudicots.
A phylogenetic study by Levin has justified the combination of Selinocarpus and Ammocodon into the genus Acleisanthes . The genus Izabalea is now considered a synonym of Agonandra , a genus in Opiliaceae. A more recent study by Douglas and Manos clarified the relationships among almost all of the genera in the family and demonstrated that a substantial diversification of herbaceous genera has occurred in arid North America. Many genera of Nyctaginaceae possess unusual characters. Notable examples include sticky bands on the stems between the nodes, cleistogamous flowers (which self-pollinate without opening), or gypsophily, the ability to grow on soils with a high concentration of gypsum.
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The extinct genus Mennegoxylon has been provisionally placed in the family. [10]
Note; as of February 2022, Plants of the World Online classes Boldoa as a synonym of Salpianthus and Grajalesia as a synonym of Pisonia. [11] [12]
The family contains one food crop, the mauka (Mirabilis extensa), a root vegetable of minor local importance in the Andes. Garden Four-O'Clocks Mirabilis jalapa species are grown as ornamental plants, as are species of Bougainvillea (Bougainvillea glabra, B. spectabilis, and numerous hybrids), Bougainvillea and Abronia are commonly cultivated in warmer regions.
The Scrophulariaceae are a family of flowering plants, commonly known as the figwort family. The plants are annual and perennial herbs, as well as shrubs. Flowers have bilateral (zygomorphic) or rarely radial (actinomorphic) symmetry. The Scrophulariaceae have a cosmopolitan distribution, with the majority found in temperate areas, including tropical mountains. The family name is based on the name of the included genus Scrophularia L.
Zygophyllaceae is a family of flowering plants that contains the bean-caper and caltrop. The family includes around 285 species in 22 genera.
The Asclepiadoideae are a subfamily of plants in the family Apocynaceae. Formerly, they were treated as a separate family under the name Asclepiadaceae, e.g. by APG II, and known as the milkweed family.
The Aizoaceae, or fig-marigold family, is a large family of dicotyledonous flowering plants containing 135 genera and about 1800 species. They are commonly known as ice plants or carpet weeds. They are often called vygies in South Africa and New Zealand. Highly succulent species that resemble stones are sometimes called mesembs.
Plantaginaceae, the plantain family, is a large, diverse family of flowering plants in the order Lamiales that includes common flowers such as snapdragon and foxglove. It is unrelated to the banana-like fruit also called "plantain." In older classifications, Plantaginaceae was the only family of the order Plantaginales, but numerous phylogenetic studies, summarized by the Angiosperm Phylogeny Group, have demonstrated that this taxon should be included within Lamiales.
Bougainvillea is a genus of thorny ornamental vines, bushes, and trees belonging to the four o' clock family, Nyctaginaceae. They are native to Brazil, Peru, and Argentina. There are between 4 and 22 species in the genus. The inflorescence consists of large colourful sepal-like bracts which surround three simple waxy flowers, gaining popularity for the plant as an ornamental. The plant is named after explorer Louis Antoine de Bougainville (1729-1811), after it was documented on one of his expeditions.
The Passifloraceae are a family of flowering plants, containing about 750 species classified in around 27 genera.
The Combretaceae, often called the white mangrove family, are a family of flowering plants in the order Myrtales. The family includes about 530 species of trees, shrubs, and lianas in ca 10 genera. The family includes the leadwood tree, Combretum imberbe. Three genera, Conocarpus, Laguncularia, and Lumnitzera, grow in mangrove habitats (mangals). The Combretaceae are widespread in the subtropics and tropics. Some members of this family produce useful construction timber, such as idigbo from Terminalia ivorensis. The commonly cultivated Quisqualis indica is now placed in the genus Combretum. Many plants in the former Quisqualis genus contain the excitotoxin quisqualic acid, a potent AMPA agonist.
The Onagraceae are a family of flowering plants known as the willowherb family or evening primrose family. They include about 650 species of herbs, shrubs, and trees in 17 genera. The family is widespread, occurring on every continent from boreal to tropical regions.
The Loganiaceae are a family of flowering plants classified in order Gentianales. The family includes up to 13 genera, distributed around the world's tropics. There are not any great morphological characteristics to distinguish these taxa from others in the order Gentianales.
Dasylirion is a genus of North American flowering succulent plants in the asparagus family, all native to Mexico and the American Southwest. In the APG III classification system, Dasylirion is placed in the family Asparagaceae, subfamily Nolinoideae.
Lomandroideae is a subfamily of monocot flowering plants in the family Asparagaceae, order Asparagales, according to the APG III system of 2009. The subfamily name is derived from the generic name of the type genus, Lomandra. The group has previously been treated as a separate family Laxmanniaceae. In the Kubitzki system, it is treated as Lomandraceae Lotsy.
Phytolaccaceae is a family of flowering plants. Though almost universally recognized by taxonomists, its circumscription has varied. It is also known as the Pokeweed family.
The Capparaceae, commonly known as the caper family, are a family of plants in the order Brassicales. As currently circumscribed, the family contains 15 genera and about 430 species. The largest genera are Capparis, Morisonia, Maerua, Boscia, and Cadaba.
Goodeniaceae is a family of flowering plants in the order Asterales. It contains about 404 species in twelve genera. The family is distributed mostly in Australia, except for the genus Scaevola, which is pantropical. Its species are found across most of Australia, being especially common in arid and semi-arid climates.
Helicteroideae is a subfamily of the family Malvaceae. Some taxonomists place genera in Helicteroideae in distinct families Durionaceae and Helicteraceae, while others recognizes the tribes Durioneae and Helictereae.
Stilbaceae is a family of flowering plants in the order Lamiales.
Montiaceae are a family of flowering plants, comprising about 14 genera with about 230 known species, ranging from small herbaceous plants to shrubs. The family has a cosmopolitan distribution.
Physaleae is a tribe of flowering plants in the subfamily Solanoideae of the family Solanaceae.
Byttnerioideae is a subfamily of the flowering plant family Malvaceae.