Effect of Carbohydrate Supplementation on Investment into Offspring Number, Size, and Condition in a Social Insect

PLoS One. 2015 Jul 21;10(7):e0132440. doi: 10.1371/journal.pone.0132440. eCollection 2015.

Abstract

Resource availability can determine an organism's investment strategies for growth and reproduction. When nutrients are limited, there are potential tradeoffs between investing into offspring number versus individual offspring size. In social insects, colony investment in offspring size and number may shift in response to colony needs and the availability of food resources. We experimentally manipulated the diet of a polymorphic ant species (Solenopsis invicta) to test how access to the carbohydrate and amino acid components of nectar resources affect colony investment in worker number, body size, size distributions, and individual percent fat mass. We reared field-collected colonies on one of four macronutrient treatment supplements: water, amino acids, carbohydrates, and amino acid and carbohydrates. Having access to carbohydrates nearly doubled colony biomass after 60 days. This increase in biomass resulted from an increase in worker number and mean worker size. Access to carbohydrates also altered worker body size distributions. Finally, we found a negative relationship between worker number and size, suggesting a tradeoff in colony investment strategies. This tradeoff was more pronounced for colonies without access to carbohydrate resources. The monopolization of plant-based resources has been implicated in the ecological success of ants. Our results shed light on a possible mechanism for this success, and also have implications for the success of introduced species. In addition to increases in colony size, our results suggest that having access to plant-based carbohydrates can also result in larger workers that may have better individual fighting ability, and that can withstand greater temperature fluctuations and periods of food deprivation.

Publication types

  • Research Support, U.S. Gov't, Non-P.H.S.

MeSH terms

  • Amino Acids / metabolism
  • Animals
  • Ants / anatomy & histology
  • Ants / growth & development
  • Ants / physiology*
  • Behavior, Animal
  • Body Size
  • Dietary Carbohydrates / metabolism*
  • Population Density
  • Reproduction

Substances

  • Amino Acids
  • Dietary Carbohydrates

Grants and funding

National Science Foundation, Division of Environmental Biology (http://www.nsf.gov/dir/index.jsp?org=BIO) 0716983 to MDE, National Science Foundation, Division of Environmental Biology (http://www.nsf.gov/dir/index.jsp?org=BIO) 0717054 to DAH, National Science Foundation, Division of Environmental Biology (http://www.nsf.gov/dir/index.jsp?org=BIO) 0716966 to AVS, and the University of Illinois, Department of Animal Biology to BDW. The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.