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}}KOS OMM AHMED HESHAMM |
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'''Black''' is a [[equine coat color|hair coat color]] of [[horse]]s in which the entire hair coat is black. Black is a relatively uncommon coat color, and it is not uncommon to mistake dark [[Chestnut (coat)|chestnuts]] or [[Bay (horse)|bays]] for black. |
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True black horses have dark brown eyes, black skin, and wholly black hair coats without any areas of permanently reddish or brownish hair. They may have pink skin beneath any [[Horse markings|white markings]] under the areas of white hair, and if such white markings include one or both eyes, the eyes may be blue. Many black horses "sun bleach" with exposure to the elements and sweat, and therefore their coats may lose some of their rich black character and may even resemble bay or [[Seal brown (horse)|seal brown]], though examination of the color of hair around the eyes, muzzle and genitals often will determine color. Black horses that do not sun bleach are called "non-fading" blacks. |
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Some [[list of horse breeds|breeds]] of horses, such as the [[Friesian horse]], [[Murgese]] and [[Ariegeois pony|Ariegeois]] (or Merens) are almost exclusively black. Black is also common in the [[Fell pony]], [[Dales pony]], [[Ostfriesen and Alt-Oldenburger]], [[Kladruber]], and [[Groningen Horse|Groningen]]. |
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==Visual identification== |
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[[Image:Brownvblack.JPG|left|thumb|Black horse (top) with sun bleached mane compared against dark bay or seal brown horse (bottom) with reddish hairs around the eye.]] |
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When identifying the base color of a horse, it is important to disregard all pink-skinned white [[Horse markings|markings]]. White markings and patterns such as [[Pinto horse|pinto]] and [[Leopard (pattern)|leopard]] have no bearing on the underlying base coat color of the animal. |
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Black [[foal]]s are typically born a mousy gray but can be darker shades. As many foals have [[primitive markings]] at birth, some black foals are mistaken for [[grullo]] or even bay dun; the primitive markings on a black foal will, however, disappear as the black hair coat grows in. Black foals have dark skin and eyes at birth. An adult-like black foal coat often indicates that the foal will [[Gray (horse)|gray]], if the foal has at least one gray parent. Graying can be confirmed by the presence of white hairs around the eyes and muzzle. Gray [[Lipizzan]]er horses are frequently born black. |
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[[File:Dales Pony Mare & Foal at the Dales Pony Society Breed Show.jpg|thumb|As seen in this photo, black foals are often born a "mousy" color that sheds off as they get older]] |
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Black adult horses are easier to identify, as the coat must be entirely black, even if superficially sun bleached. A sun bleached black may be confused with a dark [[Bay (horse)|bay]], but a trained eye can distinguish between them, particularly by examining the fine hairs around the eyes and muzzle. When a black horse is sun-bleached, the mane and tail often sun bleach most prominently, and the rest of the coat may have a rusty tinge. A sun-bleached black may also be mistaken for the less common [[smoky black]], but can be distinguished by pedigree analysis or DNA testing. |
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==Black mimics== |
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[[Image:Killesberg Shetlandfohlen.jpg|thumb|This black [[Shetland Pony]] foal was born very dark and will likely gray like its mother]] |
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* [[Bay (horse)|Dark bay]] or [[Seal brown (horse)|seal brown]]: The darkest shades of bay are commonly confused with black, even by experienced horse persons. However, a dark bay will always show some rich red character in its coat. Horses with a very dark coat that may appear black, but have tan or reddish hairs around the eyes, muzzle, armpits, and stifle are sometimes called "seal brown", "mahogany bay", or "black bay." Both colors are genetically distinct from black and can be confirmed with a [[DNA]] test. |
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* [[Chestnut (coat)|Liver chestnut]]: Some red horses are so dark that they appear black, and are often called "black chestnuts" as a consequence. However, even the darkest liver chestnuts will show some red character in their coats, usually in the hair around the pastern or in the mane or tail. Even dark liver chestnuts do not have any true black pigment in their coats. This can be verified with DNA testing. Liver chestnut is very common in the [[Morgan horse]]. |
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* [[Smoky black]]: The action of the [[cream gene]] in the [[heterozygous]] condition has a minimal effect on black pigment, so heterozygous creams with a black base coat (smoky blacks) differ little from true blacks. A smoky black will have at least one cream parent, is often born a pewter shade with blue eyes, and usually retains reddish hair inside the ear through adulthood. |
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==Genetic identification== |
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{{Main|Equine coat color genetics}} |
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[[Image:Irish Tinker horse 2.JPG|thumb|A horse with a black base coat overlain by [[tobiano]]-patterned white markings, called a [[piebald]] in some countries.]] |
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In the study and discussion of [[equine coat color genetics]], black is considered a "base" color, as is [[Chestnut (coat)|red]]. This designation makes the effects of other coat color [[gene]]s easier to understand. Coat colors that are designated "black-based" include [[grullo]] (also called blue [[dun gene|dun]]), [[smoky black]], [[smoky cream]], [[Silver dapple gene|silver black]], [[Champagne gene|classic champagne]], and [[Roan (horse)|blue roan]]. Sometimes this designation includes the bay family: [[Bay (horse)|bay]], [[Seal brown (horse)|seal brown]], [[Buckskin (horse)|buckskin]], [[Dun gene|bay dun]], [[Silver dapple gene|silver bay]], [[perlino]], [[Champagne gene|amber champagne]], and [[Roan (horse)|bay roan]]. Horses with a black-based coat may also have added spotting patterns including [[leopard (pattern)|leopard]] patterns seen on [[Appaloosa]]s and the [[pinto horse|pinto]] coloring known as [[piebald]]. |
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The genetics behind the black horse are relatively simple. The color black is primarily controlled by two genes: [[Melanocortin 1 receptor|Extension]] and [[Agouti signalling peptide|Agouti]]. The functional, [[Dominance relationship|dominant]] [[allele]] of the extension gene (labeled "E") enables the horse to produce [[eumelanin|black pigment]] in the hair. Without this gene ([[homozygous]] recessive condition "ee"), the coat is devoid of black pigment and the horse is some shade of [[Chestnut (coat)|red]]. The functional, dominant allele (or alleles) of the [[agouti gene]] (labeled "A") enable the horse to restrict black pigment to certain parts of the coat, notably the legs, mane and tail, allowing the underlying red to show through, resulting in bay coloring. Without this gene (homozygous recessive condition "aa"), any black pigment present is unrestricted, resulting in a uniformly black coat. |
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Thus a black horse has at least one copy of the functional, dominant "E" allele and two copies of the non-functional, recessive "a" allele. A mature true black horse can be safely said to possess at least one dominant extension gene (EE or Ee); and has no other dominant genes (such as agouti, gray, or any of the dilution factors) that further modify color. |
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A [[DNA test]], which uses hair with the root intact, has been developed to test for the Extension and Agouti [[genotype]]s. However, the terminology can be manipulated. Unfortunately, the extension test is often mislabeled as the "black test", leading to confusion. Neither the extension test nor the agouti test alone can identify a black horse. Together, they can determine that a horse that appears visually black is not actually a dark bay or liver chestnut. |
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Horses described as "homozygous black" are simply homozygous for the dominant extension gene (EE); they are homozygous "not-red". Such horses are only "guaranteed" to never produce a [[Chestnut (coat)|red]] foal. The actual horse may carry additional genetic modifiers that could make it bay, buckskin, gray, bay roan, perlino, silver bay, and so on. A visually black horse that is tested "homozygous black" is EE and has no other color modifiers. |
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However, it has become popular for individuals owning a horse that is homozygous for the extension gene (EE) to claim that the horse will "throw black." But, generally speaking, one horse cannot be guaranteed to "throw black" with all mates. The mate of a true black horse may contribute the a dominant Agouti allele, which will suppress the black coloring and result in a bay foal. If a black is bred to a gray, the ensuing foal may also be gray. Other modifiers present in the mate may produce additional dilution colors or spotting patterns. Nonetheless certain individual pairings with appropriate DNA testing can, in some cases, be guaranteed to produce black. |
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==References== |
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<References/> |
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*[http://www.vgl.ucdavis.edu/services/coatcolorhorse.php "Horse coat color tests"] from the [[UC Davis]] Veterinary Genetics Lab |
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*[http://www.vgl.ucdavis.edu/services/coatcolor.php "Introduction to Coat Color Genetics"] ''from'' Veterinary Genetics Laboratory, School of Veterinary Medicine, University of California, Davis. Web Site accessed January 12, 2008. |
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*[http://www.biomedcentral.com/content/pdf/1297-9686-29-3-403.pdf (pdf) D.P Sponenberg and M.C. Weise, "Dominant black in horses" in ''Genetics Selection Evolution'', Department of Biosciences and Pathobiology, Virginia-Maryland Regional College of Veterinary Medicine, no 29, mars 1997, p. 403-408] |
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==External links== |
==External links== |
Revision as of 08:39, 22 January 2020
This article includes a list of references, related reading, or external links, but its sources remain unclear because it lacks inline citations. (October 2016) |
Black | |
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Variants | Fading, non-fading, possibly genetic |
Genotype | |
Base color | Extension "E" |
Modifying genes | none |
Description | Solid black base color uniform over entire body other than markings |
Skin | Black |
Eyes | Brown |
KOS OMM AHMED HESHAMM
genetic tests for the coat colors chestnut, bay and black. Results from a preliminary study in the Swiss Freiberger horse breed", in Schweiz Arch Tierheilkd, Institut für Nutztierwissenschaften der ETH Zürich, n°29, 2002, p. 144(8):405-12
External links
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