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'''Caucasus hunter-gatherer''' ('''CHG'''), also called '''Satsurblia cluster''',<ref>{{Cite journal |last=Fu |first=Qiaomei |last2=Posth |first2=Cosimo |last3=Hajdinjak |first3=Mateja |last4=Petr |first4=Martin |last5=Mallick |first5=Swapan |last6=Fernandes |first6=Daniel |last7=Furtwängler |first7=Anja |last8=Haak |first8=Wolfgang |last9=Meyer |first9=Matthias |last10=Mittnik |first10=Alissa |last11=Nickel |first11=Birgit |last12=Peltzer |first12=Alexander |last13=Rohland |first13=Nadin |last14=Slon |first14=Viviane |last15=Talamo |first15=Sahra |date=2016-06 |title=The genetic history of Ice Age Europe |url=https://www.nature.com/articles/nature17993 |journal=Nature |language=en |volume=534 |issue=7606 |pages=200–205 |doi=10.1038/nature17993 |issn=1476-4687 |via=Nature}}</ref><ref>{{cite journal |author1=Eisenmann, S. |author2= Bánffy, E. |author3=van Dommelen, P. |display-authors=etal |year=2018 |title=Reconciling material cultures in archaeology with genetic data: The nomenclature of clusters emerging from archaeogenomic analysis |journal=Scientific Reports |volume=8 |issue= 1 |pages= 13003 |doi=10.1038/s41598-018-31123-z |pmid= 30158639 |pmc= 6115390 |bibcode= 2018NatSR...813003E }}</ref> is an [[anatomically modern human]] [[human genetics|genetic lineage]], first identified in a 2015 study,{{sfn|Jones|Gonzalez-Fortes|Connell|Siska|2015}}{{sfn|Fu|Posth|Hajdinjak|Petr|2016}} based on the [[population genetics]] of several modern [[Eurasia|Western Eurasian]] ([[Europe]]an, [[Caucasus|Caucasian]] and [[Near Eastern]]) populations.<ref name="bbcnov16"/><ref name="Dutchen 2016">{{cite web|last1=Dutchen|first1=Stephanie|title=History on Ice|url=http://hms.harvard.edu/news/history-ice|publisher=[[Harvard Medical School]]|access-date=11 May 2016|date=2 May 2016}}</ref>
It represents an ancestry maximised in some [[Upper Paleolithic]] and [[Mesolithic]] hunter-gatherer groups in the [[Caucasus]]. These groups are also very closely related to Mesolithic hunter-gatherers and [[Neolithic]] farmers in the [[Iranian Plateau]], who are sometimes included within the CHG group.<ref>{{Cite journal |last=Lazaridis |first=Iosif |last2=Alpaslan-Roodenberg |first2=Songül |last3=Acar |first3=Ayşe |last4=Açıkkol |first4=Ayşen |last5=Agelarakis |first5=Anagnostis |last6=Aghikyan |first6=Levon |last7=Akyüz |first7=Uğur |last8=Andreeva |first8=Desislava |last9=Andrijašević |first9=Gojko |last10=Antonović |first10=Dragana |last11=Armit |first11=Ian |last12=Atmaca |first12=Alper |last13=Avetisyan |first13=Pavel |last14=Aytek |first14=Ahmet İhsan |last15=Bacvarov |first15=Krum |date=26 August 2022 |title=The genetic history of the Southern Arc: A bridge between West Asia and Europe |url=https://www.science.org/doi/10.1126/science.abm4247 |journal=Science |language=en |volume=377 |issue=6609 |doi=10.1126/science.abm4247 |issn=0036-8075 |pmc=10064553 |pmid=36007055}}</ref><ref>{{Cite journal |last=Wang |first=Chuan-Chao |last2=Reinhold |first2=Sabine |last3=Kalmykov |first3=Alexey |last4=Wissgott |first4=Antje |last5=Brandt |first5=Guido |last6=Jeong |first6=Choongwon |last7=Cheronet |first7=Olivia |last8=Ferry |first8=Matthew |last9=Harney |first9=Eadaoin |last10=Keating |first10=Denise |last11=Mallick |first11=Swapan |last12=Rohland |first12=Nadin |last13=Stewardson |first13=Kristin |last14=Kantorovich |first14=Anatoly R. |last15=Maslov |first15=Vladimir E. |date=4 February 2019 |title=Ancient human genome-wide data from a 3000-year interval in the Caucasus corresponds with eco-geographic regions |url=https://www.nature.com/articles/s41467-018-08220-8 |journal=Nature Communications |language=en |volume=10 |issue=1 |doi=10.1038/s41467-018-08220-8 |issn=2041-1723 |pmc=6360191 |pmid=30713341}}</ref> Ancestry that is closely related to CHG-Iranian Neolithic farmers is also known from further east, including from the [[Bactria–Margiana Archaeological Complex]] and the [[Indus Valley Civilisation|Harappan/Indus Valley Civilisation]].<ref name=":2">{{Cite journal |last=Shinde |first=Vasant |last2=Narasimhan |first2=Vagheesh M. |last3=Rohland |first3=Nadin |last4=Mallick |first4=Swapan |last5=Mah |first5=Matthew |last6=Lipson |first6=Mark |last7=Nakatsuka |first7=Nathan |last8=Adamski |first8=Nicole |last9=Broomandkhoshbacht |first9=Nasreen |last10=Ferry |first10=Matthew |last11=Lawson |first11=Ann Marie |last12=Michel |first12=Megan |last13=Oppenheimer |first13=Jonas |last14=Stewardson |first14=Kristin |last15=Jadhav |first15=Nilesh |date=October 2019 |title=An Ancient Harappan Genome Lacks Ancestry from Steppe Pastoralists or Iranian Farmers |url=https://linkinghub.elsevier.com/retrieve/pii/S0092867419309675 |journal=Cell |language=en |volume=179 |issue=3 |pages=729–735.e10 |doi=10.1016/j.cell.2019.08.048 |pmc=6800651 |pmid=31495572}}</ref><ref name=":3">{{Cite journal |last=Narasimhan |first=Vagheesh M. |last2=Patterson |first2=Nick |last3=Moorjani |first3=Priya |last4=Rohland |first4=Nadin |last5=Bernardos |first5=Rebecca |last6=Mallick |first6=Swapan |last7=Lazaridis |first7=Iosif |last8=Nakatsuka |first8=Nathan |last9=Olalde |first9=Iñigo |last10=Lipson |first10=Mark |last11=Kim |first11=Alexander M. |last12=Olivieri |first12=Luca M. |last13=Coppa |first13=Alfredo |last14=Vidale |first14=Massimo |last15=Mallory |first15=James |date=6 September 2019 |title=The formation of human populations in South and Central Asia |url=https://www.science.org/doi/10.1126/science.aat7487 |journal=Science |language=en |volume=365 |issue=6457 |doi=10.1126/science.aat7487 |issn=0036-8075 |pmc=6822619 |pmid=31488661}}</ref> Caucasus hunter-gatherers and [[Eastern hunter-gatherer]]s are ancestral in roughly equal proportions to the [[Western Steppe Herder]]s (WSH), who were widely spread across Europe and Asia beginning during the [[Chalcolithic]].
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