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'''Plotopteridae'''<ref>{{cite journal|author= Howard, H. |year=1969|title= A new avian fossil from Kern County, California|journal=[[Condor (journal)|Condor]]|volume=71|issue=1|pages= 68–69|url=http://sora.unm.edu/sites/default/files/journals/condor/v071n01/p0068-p0069.pdf|doi= 10.2307/1366050|jstor=1366050}}</ref> is the name of an [[extinct]] [[family (biology)|family]] of flightless [[seabird]]s from the order [[Suliformes]]. Related to the [[gannet]]s and [[booby|boobies]], they exhibited remarkable [[convergent evolution]] with the [[penguin]]s, particularly with the now [[extinct]] giant penguins.<ref>{{cite journal|author= Olson, Storrs L.
Their [[fossil]]s have been found in [[California]], [[Oregon]],<ref>{{cite journal|author= Goedert, James L. |year=1988| title= A new late Eocene species of Plototpteridae (Aves: Pelecaniformes) from northwestern Oregon |journal= Proceedings of the California Academy of Sciences |volume= 45 |pages= 97–102}}</ref> [[Washington (U.S. state)|Washington]],<ref>{{Cite journal|author1=Gerald Mayr |author2=James L. Goedert |lastauthoramp=yes |year=2016 |title=New late Eocene and Oligocene remains of the flightless, penguin-like plotopterids (Aves, Plotopteridae) from western Washington State, U.S.A. |journal=Journal of Vertebrate Paleontology |volume=36 |issue=4 |pages=e1163573 |doi=10.1080/02724634.2016.1163573 }}</ref><ref>{{cite journal| author1= Mayr, Gerald| author2= Goedert, James L.| year= 2018| title= First record of a tarsometatarsus of ''Tonsala hildegardae'' (Plotopteridae) and other avian remains from the late Eocene/early Oligocene of Washington State (USA)| journal= Geobios| volume= 51| pages= 51–59| doi= 10.1016/j.geobios.2017.12.006 }}</ref> [[British Columbia]]<ref>{{Cite journal|author1=Gary Kaiser
[[File:Tonsala hildegardae.png|thumb|left|''Tonsala hildegardae'' fossils]]
The second species to be named from rocks along the eastern Pacific Ocean was ''Tonsala hildegardae''<ref>{{Cite journal| author= Olson, Storrs L.| year= 1980| title= A new genus of penguin-like pelecaniform bird from the Oligocene of Washington (Pelecaniformes: Plotopteridae)| journal= Contributions in Science| volume= 330| pages= 51–57 }}</ref> from the late Oligocene lower part of the Pysht Formation in Washington State. More fossils of ''T. hildegardae'' have since been described <ref>{{Cite journal| author1 = Goedert, James L.| author2= Cornish, John| year= 2002| title= A preliminary report on the diversity and stratigraphic distribution of the Plotopteridae (Pelecaniformes) in Paleogene rocks of Washington State, USA| journal= Proceedings of the 5th Symposium of the Society of Avian Paleontology and Evolution, Beijing| pages= 63–76}}</ref><ref>{{cite journal| author1= Mayr, Gerald| author2= Goedert, James L.| year= 2017| title= First record of a tarsometatarsus of ''Tonsala hildegardae'' (Plotopteridae) and other avian remains from the late Eocene/early Oligocene of Washington State (USA)| journal= Geobios| volume= 51| pages= 51–59}}</ref> and included some of the first known examples of borings made by the marine bone-eating worm ''Osedax'' in bird bones.<ref>{{Cite journal| author1= Kiel, Steffen| author2= Kahl, W.-A.| author3= Goedert, James L.| year= 2011| title= ''Osedax'' borings in fossil marine bird bones| journal= Naturwissenschaften| volume= 98| pages= 51–55}}</ref>
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{{Reflist}}
*{{cite journal|author= Howard, H. |year=1969|title= A new avian fossil from Kern County, California|journal=[[Condor (journal)|Condor]]|volume=71|issue=1|pages= 68–69|url=http://sora.unm.edu/sites/default/files/journals/condor/v071n01/p0068-p0069.pdf|doi= 10.2307/1366050|jstor=1366050}}
*{{cite journal|author= Olson, Storrs L.
*{{cite journal|author1=Olson, Storrs L. |author2=Hasegawa, Yoshikazu |lastauthoramp=yes |year=1996|title= A new genus and two new species of gigantic Plotopteridae from Japan (Aves: Pelecaniformes)|journal=[[Journal of Vertebrate Paleontology|J. Vert. Paleontol.]]|volume=16|issue=4|pages= 742–751|doi= 10.1080/02724634.1996.10011362}}
*{{cite book|author1=Schreiber, E.A. |author2=Burger, J. |lastauthoramp=yes |year=2001|title= Biology of Marine Birds|isbn=0-8493-9882-7}}
*{{cite journal|author= Mayr, Gerald |year=2005|title= Tertiary plotopterids (Aves, Plotopteridae) and a novel hypothesis on the phylogenetic relationships of penguins (Spheniscidae)|journal=Journal of Zoological Systematics|volume=43|issue=1|pages= 67–71|url=http://www.senckenberg.de/files/content/forschung/abteilung/terrzool/ornithologie/plotopteridae.pdf|doi=10.1111/j.1439-0469.2004.00291.x}}
*{{Cite journal|author1=Gary Kaiser
*{{Cite journal|author1=Gerald Mayr |author2=James L. Goedert |lastauthoramp=yes |year=2016 |title=New late Eocene and Oligocene remains of the flightless, penguin-like plotopterids (Aves, Plotopteridae) from western Washington State, U.S.A. |journal=Journal of Vertebrate Paleontology |volume=36 |issue=4 |pages=e1163573 |doi=10.1080/02724634.2016.1163573 }}
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